Knuckle was a slaughterer and self-taught pilot who flew a skycraft called the Woodwasp in the Silver Pastures and the Free Glades. He rescued Rook from certain death during an attack from a logworm during the events of The Last of the Sky Pirates.

A year later, he assisted Varis Lodd and Rook during a raid on the Foundry Glades, in which they attempted to rescue a group enslaved Banderbears.

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